MAIN PATHWAYS TO AND FROM THE CEREBELLUM:
THE VESTIBULAR SYSTEM
THE INFERIOR OLIVE - CEREBELLUM CONNECTIONS:
Climbing Fiber Interconnections via the Inferior Olive:
CEREBRAL PATHWAYS TO AND FROM THE CEREBELLUM:
The unique structure of the cerebellum is shown to integrate spinal and cranial sensory inputs with cerebral processes and
motor outputs, synapse by synapse. Focusing primarily on the vestibular system alternate regenerative and expressive
modes of System 4 are shown to meaningfully span and integrate events in space and time. (Very concentrated focus is
required to grasp the interrelated complexity involved.)
Review of System 4 Transformations as Outlined in Part 1- Spinal Cord:
The Nervous System - Part 1 – Spinal Integration reviewed System 4 briefly. There are three Particular Sets of active energy
interfaces that transform through a Six Step Sequence of different Terms, each set transforming through the same Term
sequence one Step apart. Particular Terms also have Expressive and Regenerative Modes that interact in the synchronous
matrix of transformations that is regulated and integrated by two Universal Sets in recurrent four step Cycles. There are thus
12 Steps needed to complete both an expressive and a regenerative sequence.
Chart Summary of Twelve Step Sequence:
The following chart from Part 1 is reproduced here for easier reference. Expressive and regenerative particular terms, as
well as universal terms, are shown for each sequential Step. Regenerative Terms are shown in bold. For an explanation of
how the Universal Terms work see Part 1.
Keep in mind that in general each System 4 Step corresponds to a synapse in the nervous system.
Step Set 1 Set 2 Set 3 Set U1 Set U2 Cycle
1 T8E T7R T4E T9 T3 #1
2 T5E T1R T2E T9 T6
3 T7E T4R T8E T8R T6
4 T1E T2R T5R T8R T2E
5 T4E T8E T7R T9 T3 #2
6 T2E T5E T1R T9 T6
7 T8E T7E T4R T8R T6
8 T5R T1E T2R T8R T2E
9 T7R T4E T8E T9 T3 #3
10 T1R T2E T5E T9 T6
11 T4R T8E T7E T8R T6
12 T2R T5R T1E T8R T2E
|The Human Nervous System - Part 2 - The Cerebellum
By Robert Campbell 2006
In Step 4 of this previous cycle there is an initial assessment of the preparedness of hair cell and
muscle action potentials and tone. This can affect the selection of the pattern of cell activation
that follows in the next Cycle 1 below.
Nervous System - Part 2 - The Cerebellum
Hair cells activate primary projections of the vestibular system. Muscle spindles monitor parent
muscle action and project to relay nuclei in the Central Nervous System (CNS).
Primary neurons project from hair cells to vestibular nuclei as Idea Terms. Proprioceptive relay
nuclei in the CNS also project as Idea Terms to the vestibular nuclei from all spinal levels
Parallel patterned response outputs that balance sensory vestibular inputs project to eye, neck,
and spinal motor nuclei.
The motor output to eye & neck muscles is reconciled with other inputs to these motor nuclei.
The eyes may move to compensate for head movement or move separately. This output is
regenerative since it is governed by connections established through prior learning of the
individual and the species. The motor output to the neck and eyes is via the Medial Longitudinal
Fasciculus (MLF). The motor output to the ventral motor horns of the spinal column is via the
lateral vestibulospinal tract.
Memory of the sequence is stored in dendritic and synaptic protein synthesis of cells involved, as
well as the Void. The synaptic connections evolve consistent with learning experience. At the
same time related regenerative memories are spontaneously recalled in the reticular formation.
This activates efferent projections to hair cells and muscle spindles.
The regenerative efferent projections to hair cells and muscle spindles initiate a patterned motor
simulation. In the projection to muscle spindles this takes place in gamma motor neurons that
project to the spindles.
An actual simulation takes place in hair cells and in neck and body muscle spindles. The hair cell
simulation involves the active movement of the kinocilium in the Type 2 hair cell with respect to
the current state of the Type 1 hair cells. The muscle spindle simulation likewise simulates a
pattern of movement with respect to the current position of muscles.
The simulations project as regenerative vestibular input and likewise corrective proprioceptive
feedback to vestibular nuclei as a regenerative idea to restore balance.
The simulations result in a synapse in the vestibular nuclei to a pattern of planned motor output
to eye, neck and body muscles via the MLF and the lateral vestibulospinal tracts as in the T8E
Term above. All Particular Set T8 Terms are expressive. (See Part 1) .
The output of vestibular nuclei to eye, neck and body spinal motor nuclei for corrective action is
reconciled with other inputs to these nuclei as they project to muscles. This output is expressive
since it is creatively planned by proprioceptive input. It is patterned action of the Host(1) that
directs Organs(2) whereas in the regenerative sequence it is conditioned Organ(2) processes
established through experience that direct the Host(1) . (See Part 1)
Memory of the action sequence is stored and expressive memories recalled similar to T7R above.
There is a perception of the capacity to respond to ongoing vestibular needs. This involves the
recovery of cells processes such as action potentials and the stabilization of fluid in semi-circular
canals, preparatory for the next action sequence.